Betacoronavirus gravedinis | |
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Transmission electron micrograph of human coronavirus OC43 | |
Virus classification | |
(unranked): | Virus |
Realm: | Riboviria |
Kingdom: | Orthornavirae |
Phylum: | Pisuviricota |
Class: | Pisoniviricetes |
Order: | Nidovirales |
Family: | Coronaviridae |
Genus: | Betacoronavirus |
Subgenus: | Embecovirus |
Species: | Betacoronavirus gravedinis
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Member viruses[1] | |
Synonyms | |
Betacoronavirus gravedinis [4] (also called Betacoronavirus 1) is a species of coronavirus which infects humans and cattle.[5][6] The infecting virus is an enveloped, positive-sense, single-stranded RNA virus and is a member of the genus Betacoronavirus and subgenus Embecovirus.[7] Like other embecoviruses, it has an additional shorter spike-like surface protein called hemagglutinin esterase (HE) as well as the larger coronavirus spike protein.[8][9]
A new species, Betacoronavirus 1, is proposed to accommodate previously established Betacoronavirus species of humans, cattle, swine and horses (Human enteric coronavirus; Human coronavirus OC43; Bovine coronavirus; Porcine hemagglutinating encephalomyelitis virus; Equine coronavirus) and a newly recognized virus of dogs (canine respiratory coronavirus; Erles et al., 2007)
See Table 1.
In all members of Betacoronavirus subgroup A, a haemagglutinin esterase (HE) gene, which encodes a glycoprotein with neuraminate O-acetyl-esterase activity and the active site FGDS, is present downstream to ORF1ab and upstream to S gene (Figure 1).
The presence of HE genes exclusively in members of Betacoronavirus subgroup A, but not members of Betacoronavirus subgroup B, C and D suggested that the recombination had probably occurred in the ancestor of members of Betacoronavirus subgroup A, after diverging from the ancestor of other subgroups of Betacoronavirus.