Columella (gastropod)

The columella (meaning "little column") or (in older texts) pillar is a central anatomical feature of a coiled snail shell, a gastropod shell. The columella is often only clearly visible as a structure when the shell is broken, sliced in half vertically, or viewed as an X-ray image.

The columella runs from the apex of the shell to the midpoint of the undersurface of the shell, or the tip of the siphonal canal in those shells which have a siphonal canal. If a snail shell is visualized as a cone of shelly material which is wrapped around a central axis, then the columella more or less coincides spatially with the central axis of the shell. In the case of shells that have an umbilicus, the columella is a hollow structure.

The columella of some groups of gastropod shells can have a number of plications or folds (the columellar fold, plaits or plicae), which are usually visible when looking to the inner lip into the aperture of the shell. These folds can be wide or narrow, prominent or subtle. These features of the columella are often useful in identifying the family, genus, or species of the gastropod.

The surface of the columella is called the columellar wall. The columellar callus is a smooth, calcareous thickening, secreted by the mantle, extending over the columellar area. The columellar lip, the visible part of the columella, is the lower part of the inner lip and is situated near the axis of coiling. A columellar tooth is a raised projection on the inner lip of a columella in the direction of the aperture.