The mRNA decapping complex is a protein complex in eukaryotic cells responsible for removal of the 5' cap.[1] The active enzyme of the decapping complex is the bilobed Nudix family enzyme Dcp2, which hydrolyzes 5' cap and releases 7mGDP and a 5'-monophosphorylated mRNA.[1] This decapped mRNA is inhibited for translation and will be degraded by exonucleases.[2] The core decapping complex is conserved in eukaryotes. Dcp2 is activated by Decapping Protein 1 (Dcp1) and in higher eukaryotes joined by the scaffold protein VCS.[3] Together with many other accessory proteins, the decapping complex assembles in P-bodies in the cytoplasm.