Dromasauria

For the group of dinosaurs, see Dromaeosauridae.

"Dromasauria"
Temporal range: Middle Permian - Late Permian
Life restoration of Galechirus
Life restoration of Galechirus
Scientific classificationEdit this classification
Domain: Eukaryota
Kingdom: Animalia
Phylum: Chordata
Clade: Synapsida
Clade: Therapsida
Suborder: †Anomodontia
Informal group: †Dromasauria
Broom, 1907
Included genera

"Dromasaurs" are an artificial grouping of small anomodont therapsids from the Middle and Late Permian of South Africa. They represent either a paraphyletic grade or a polyphyletic grouping of small non-dicynodont basal anomodonts rather than a clade, and as such are considered an invalid group today. "Dromasaurs" were historically united by their superficially similar appearances that were unlike other known anomodonts. They are all small in size with slender limbs and long tails, and have short skulls with very large eye sockets. "Dromasauria" (sometimes also known as "Dromasauroidea") traditionally includes three genera, all from the Karoo Supergroup of South Africa: Galepus, Galechirus, and Galeops. These genera have sometimes been divided into two subgroups, the monotypic family Galeopidae (containing only Galeops) and the Galechiridae for Galechiris and Galepus.[1]

Despite their superficial similarities, "Dromasauria" is not recognised in modern cladistics-based taxonomy (where groups are based upon shared common ancestry). Rather than forming their own clade, phylogenetic analyses have found the various "dromasaurs" to be distributed individually throughout the evolutionary tree of basal anomodonts. In particular, Galeops is notably found to consistently be much closer to the dicynodonts than to the other "dromasaurs". Some earlier studies have inferred or even recovered a close sister-relationship between Galechirus and Galepus in a clade, to which the name Galechiridae has sometimes been applied.[2][3] However, more recent phylogenetic analyses incorporating more data and more complete samples of basal anomodonts have found them at separate points on the tree too.[4]

The cladogram below depicts the results of the phylogenetic analysis from Angielczyk and Kammerer (2017), with each of the three "dromasaurs" highlighted in light green:[4]

Anomodontia    "Dromasaurs"
Galepus.
  1. ^ Broom, R. (1912). "On some new Fossil Reptiles from the Permian and Triassic Beds of South Africa". Proceedings of the Zoological Society of London. 84 (4): 859–876. doi:10.1111/j.1469-7998.1912.tb07564.x.
  2. ^ Rubidge, B. S.; Hopson, J. A. (1996). "A primitive anomodont therapsid from the base of the Beaufort Group (Upper Permian) of South Africa". Zoological Journal of the Linnean Society. 117 (2): 115–139. doi:10.1111/j.1096-3642.1996.tb02152.x.
  3. ^ Kammerer, C. F.; Angielczyk, K. D.; Fröbisch, J. (2011). "A comprehensive taxonomic revision of Dicynodon (Therapsida, Anomodontia) and its implications for dicynodont phylogeny, biogeography, and biostratigraphy". Journal of Vertebrate Paleontology. 31 (Suppl. 1): 1–158. doi:10.1080/02724634.2011.627074. S2CID 84987497.
  4. ^ a b Angielczyk, K. D.; Kammerer, C. F. (2017). "The cranial morphology, phylogenetic position and biogeography of the upper Permian dicynodont Compsodon helmoedi van Hoepen (Therapsida, Anomodontia)". Papers in Palaeontology. 3 (4): 513–545. doi:10.1002/spp2.1087. S2CID 134092461.