| Born | c. 1945 Amboseli National Park, Kenya |
|---|---|
| Died | May 3, 2009 (aged 63–64) |
Echo was an African bush elephant matriarch who was studied for over 30 years by ethologist Cynthia Moss, beginning in 1973,[1] and was the subject of several books and films. She was the first subject of the Amboseli Elephant Research Project, the longest-running study of a land mammal. The study of Echo and her family contributed significantly to the understanding of elephants, including their life-cycles, methods of communication, emotional lives, and cooperative care of the young.[2]
Echo died on May 3, 2009, around 63 or 64 years of age.[2]