Glyptodont

Glyptodonts Temporal range: Eocene–Late Pleistocene (Divisaderan–Lujanian) ~38–0.011 Ma PreꞒ Ꞓ O S D C P T J K Pg N
Glyptodon fossil, Natural History Museum, Vienna
Illustration of the skeleton of Doedicurus
Scientific classification
Domain:	Eukaryota
Kingdom:	Animalia
Phylum:	Chordata
Class:	Mammalia
Order:	Cingulata
Family:	Chlamyphoridae
Subfamily:	†Glyptodontinae Burmeister 1879
Genera
†Andinoglyptodon †Asterostemma †Boreostemma †Castellanosia †Clypeotherium †Cranithlastus †Daedicuroides †Doedicurus †Eleutherocercus †Eucinepeltus †Glyptatelus †Glyptodon †Glyptodontidium †Glyptostracon †Glyptotherium †Hoplophorus †Kelenkura †Lomaphorus †Metopotoxus †Neosclerocalyptus †Neuryurus †Palaehoplophorus †Panochthus †Paraeucinepeltus †Parapropalaehoplophorus †Plaxhaplous †Prodaedicurus †Propalaehoplophorus †Protoglyptodon †Pseudoplohophorus †Stromatherium †Xiphuroides

Glyptodonts are an extinct clade of large, heavily armoured armadillos, reaching up to 1.5 metres (4.9 ft) in height, and maximum body masses of around 2 tonnes. They had short, deep skulls, a fused vertebral column, and a large bony carapace made up of hundreds of individual scutes. Some glyptodonts had clubbed tails, similar to ankylosaurid dinosaurs.^[1]

The earliest widely recognised fossils of glyptodonts in South America are known from the late Eocene, around 38 million years ago, and they spread to southern North America after the continents became connected around 2.7 million years ago.^[2] The best-known genus within the group is Glyptodon.

Glyptodonts were historically considered to constitute the distinct family Glyptodontidae, with their relationships to modern armadillos being contested. In 2016, an analysis of the mitochondrial genome of Doedicurus found that it was, in fact, nested within the modern armadillos as the sister group of a clade consisting of Chlamyphorinae (fairy armadillos) and Tolypeutinae (giant, three-banded and naked-tailed armadillos). For this reason, glyptodonts and all armadillos but Dasypus (long-nosed or naked-tailed armadillos) were relocated to a new family, Chlamyphoridae, and glyptodonts were demoted to the subfamily Glyptodontinae.^[2][1] Other authors have continued to use Glyptodontidae.^[3] Based on the morphology of the inner ear, a close relationship with pampatheres has also been proposed.^[4]

Glyptodonts abruptly became extinct approximately 12,000 years ago at the end of the Late Pleistocene, as part of the end-Pleistocene extinction event, along with most other large animals in the Americas. Evidence has been found suggesting that they were hunted by recently arrived Paleoindians, which may have played a role in their extinction.^[3]