Hadrianus | |
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Hadrianus majusculus | |
Scientific classification | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Chordata |
Class: | Reptilia |
Order: | Testudines |
Suborder: | Cryptodira |
Clade: | Pantestudinoidea |
Superfamily: | Testudinoidea |
(unranked): | †Pantestudinidae |
Genus: | †Hadrianus Cope, 1872[2][1] |
Species | |
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Hadrianus is an extinct genus of tortoise belonging to the Testudinidae[1] found in the United States, the Yolomécatl Formation of Mexico, the Alai Beds of Kyrgyzstan and Spain[citation needed] and believed to be the oldest true tortoise known.[3] The genus is thought to be closely related to the genus Manouria.[3] The genus may have evolved in the subtropics of Asia and subsequently migrated to North America and Europe.[3] Evangelos Vlachos (2018) reassessed the North American species attributed to the genus, and determined only two as accepted namely H. corsoni & H. majusculus. The remaining species were identified as either junior synonyms, moved to other genera or considered nomen dubium do to incomplete fossils.[4]