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^Cole, C.J.; C.R. Townsend; R.P. Reynolds; R.D. MacCulloch & A. Lathrop (2013). "Amphibians and reptiles of Guyana, South America: Illustrated keys, annotated species accounts, and a biogeographic synopsis". Proceedings of the Biological Society of Washington. 125 (4): 317–620. doi:10.2988/0006-324x-125.4.317. S2CID86665287.