| Phyllolepida Temporal range: Devonian
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|---|---|
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| Austrophyllolepis ritchiei and Bothriolepis mawsoni | |
Scientific classification 
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| Domain: | Eukaryota |
| Kingdom: | Animalia |
| Phylum: | Chordata |
| Class: | †Placodermi |
| Order: | †Arthrodira |
| Suborder: | †Actinolepidoidei |
| Clade: | †Phyllolepida Stensiö 1934 |
| Type species | |
| Phyllolepis concentrica Agassiz 1844
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| Families | |
Phyllolepida ("leaf scales") is an extinct taxon of flattened placoderms found throughout the world, with fossils being found in Devonian strata. Like other flattened placoderms, the phyllolepids were bottom-dwelling predators that ambushed prey. Unlike other flattened placoderms, the phyllolepids were inhabitants of freshwater environments.
Unlike the Rhenanida, the armor of the phyllolepids were made of whole plates, rather than numerous tubercles and scales, and unlike the Petalichthyida, the components of the comparatively wide mouth are known. The phyllolepids are considered to have been blind, as the orbits for the eyes are extremely small, so much so as to suggest that the eyes were vestigial, and that they were placed on the sides of the head, as opposed to visually-oriented bottom-dwelling predators, like, say stargazers or flatfish, which have the eyes placed high on top of the head.
Despite having a relatively clear idea of the phyllolepids' lifestyle and anatomy, most fossils consist of fragments of their thoracic armor, and only two genera, Phyllolepis and Austrophyllolepis have been thoroughly studied. From the articulation of the thoracic and head plates, it has been suggested that they are either the sister group of order Arthrodira, or are in fact, a group of highly derived arthrodires.