Pioneer species

Pioneer species are resilient species that are the first to colonize barren environments, or to repopulate disrupted biodiverse steady-state ecosystems as part of ecological succession. A number of kinds of events can create good conditions for pioneers, including disruption by natural disasters, such as wildfire, flood, mudslide, lava flow or a climate-related extinction event^[1] or by anthropogenic habitat destruction, such as through land clearance for agriculture or construction or industrial damage. Pioneer species play an important role in creating soil in primary succession, and stabilizing soil and nutrients in secondary succession.^[2]

For humans, because pioneer species quickly occupy disrupted spaces they are sometimes treated as weeds or nuisance wildlife, such as the common dandelion or stinging nettle.^[3][2] Even though humans have mixed relationships with these plants, these species tend to help improve the ecosystem because they can break up compacted soils and accumulate nutrients that help with a transition back to a more mature ecosystem.^[4][5][2] In human managed ecological restoration or agroforestry, trees and herbaceous pioneers can be used to restore soil qualities and provide shelter for slower growing or more demanding plants.^[5][6][7] Some systems use introduced species to restore the ecosystem,^[2] or for environmental remediation.^[8] The durability of pioneer species can also make them potential invasive species.^{[9] [2][10]}