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Trilobites are used as index fossils to subdivide the Cambrian period. Assemblages of trilobites define trilobite zones.[1] The Olenellus-zone has traditionally marked the top of the Lower Cambrian,[2] and is followed by the Eokochaspis zone.[3]
The last two zones of the Middle Cambrian are the Bathyuriscus–Elrathina-zone (contemporaneous with the Burgess Shale) and the subsequent Bolaspidella-zone (starting at the base of the Drumian stage[4]). These are overlain by the lowermost Upper Cambrian Cedaria-zone.[5]
Alternative zoning names place the Burgess Shale in the Peronopsis bonnerensis-zone, which is underlain by the Oryctocephalus indicus-zone (e.g. Spence Shale) and overlain (perhaps not directly) by the Ptychagnostus punctuosus-zone.[6]