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The neural basis of prey detection, recognition, and orientation was studied in depth by Jörg-Peter Ewert in a series of experiments that made the toad visual system a model system in neuroethology (neural basis of natural behavior). He began by observing the natural prey catching behavior of the common European toad (Bufo bufo).
Ewert's work with toads yielded several important discoveries (Ewert 1974, 2004). In general, his research revealed the specific neural circuits for recognition of complex visual stimuli. Specifically, he identified two main regions of the brain, the tectum and the thalamic-pretectal region, that were responsible for discriminating prey from non-prey and revealed the neural pathways that connected them. Furthermore, he found that the neural mechanisms are plastic and adaptable to varying environments and conditions (Carew 2000; Zupanc 2004).