Some four-winged insect orders, such as the Lepidoptera, have developed a wide variety of morphological wing coupling mechanisms in the imago which render these taxa as "functionally dipterous" (effectively two-winged) for efficient insect flight.[1] All but the most basal forms exhibit this wing coupling.[2]: 4266
The mechanisms are of three different types - jugal, frenulo-retinacular and amplexiform.[3]