Genus of beetles
Agabus is a large genus of predatory aquatic beetles in the family Dytiscidae , proposed in 1817 by William Elford Leach [ 1] and named after Agabus , an early follower of Christianity. The adult beetles are moderate-sized, 5 to 14 mm long. The genus is primarily Holarctic in distribution, with only a few species known from the Afrotropical and Neotropical realms .[ 2] [ 3] Three species of Agabus , namely A. clypealis ,[ 4] A. discicollis [ 5] and A. hozgargantae [ 6] are endangered according to the IUCN Red List . The division into subgenera is not widely accepted. However, a number of species groups are recognized after the works of David J. Larson and Anders N. Nilsson.[ 3] The genus is probably polyphyletic or paraphyletic .[ 2] In a recent study of mitochondrial DNA , Agabus was found paraphyletic with respect to several of the species groups of Platambus , a closely related genus in the tribe Agabini.[ 7] Lately the taxonomy of the genus has been revised, and some groups of species were transferred from Agabus sensu stricto to other genera in the tribe Agabini.[ 8]
Agabus contains the following species:[ 9]
Agabus abessinicus (Zimmermann, 1928)
Agabus adpressus Aubé, 1837
Agabus adustus Guignot, 1954
Agabus aequabilis (Gschwendtner, 1923)
Agabus aequalis Sharp, 1882
Agabus aeruginosus Aubé, 1838
Agabus affinis (Paykull, 1798)
Agabus africanus Pederzani & Schizzerotto, 1998
Agabus ajax Fall, 1922
Agabus alexandrae Ribera, Hernando & Aguilera, 2001
Agabus alinae (Lafer, 1988)
Agabus ambiguus (Say, 1823)
Agabus ambulator Régimbart, 1895
Agabus amnicola (J.Sahlberg, 1880)
Agabus amoenus Solsky, 1874
Agabus ancillus Fall, 1922
Agabus angusi Nilsson, 1994
Agabus antennatus Leech, 1939
Agabus anthracinus Mannerheim, 1852
Agabus approximatus Fall, 1922
Agabus arcticus (Paykull, 1798)
Agabus aubei Perris, 1869
Agabus audeni Wallis, 1933
Agabus austinii Sharp, 1882
Agabus bakeri (Zimmermann, 1924)
Agabus basalis (Gebler, 1829)
Agabus bergi Zaitzev, 1913
Agabus bicolor (Kirby, 1837)
Agabus bifarius (Kirby, 1837)
Agabus biguttatus (Olivier, 1795)
Agabus biguttulus (Thomson, 1867)
Agabus binotatus Aubé, 1837
Agabus bipustulatus (Linnaeus , 1767)
Agabus blatta Jakovlev, 1897
Agabus brandti Harold, 1880
Agabus browni Kamiya, 1934
Agabus brunneus (Fabricius, 1798)
Agabus canadensis Fall, 1922
Agabus caraboides Sharp, 1882
Agabus cephalotes Reiche, 1861
Agabus charon Wickham, 1912
Agabus clavicornis Sharp, 1882
Agabus clypealis (Thomson, 1867)
Agabus colymbus Leech, 1938
Agabus confinis (Gyllenhal, 1808)
Agabus congener (Thunberg, 1794)
Agabus congeneroides Lomnicki , 1894
Agabus conspersus (Marsham, 1802)
Agabus conspicuus Sharp, 1873
Agabus corticeus Lomnicki, 1894
Agabus costulatus (Motschulsky, 1859)
Agabus coxalis Sharp, 1882
Agabus crassipes (Fall, 1922)
Agabus crypticoides Régimbart, 1895
Agabus debilipes Régimbart, 1899
Agabus dichrous Sharp, 1878
Agabus didymus (Olivier, 1795)
Agabus dilatatus (Brullé, 1832)
Agabus discolor (Harris, 1828)
Agabus disintegratus (Crotch, 1873)
Agabus dytiscoides Régimbart, 1908
Agabus elongatus (Gyllenhal, 1826)
Agabus erytropterus (Say, 1823)
Agabus faldermanni Zaitzev, 1927
Agabus falli (Zimmermann, 1934)
Agabus freudei Guéorguiev, 1975
Agabus friedrichi (Falkenström, 1936)
Agabus fulvaster Zaitzev, 1906
Agabus fulvipennis Régimbart, 1899
Agabus fuscipennis (Paykull, 1798)
Agabus galamensis Nilsson, 1992
Agabus glacialis Hochhuth, 1846
Agabus glazunovi (Zaitzev, 1953)
Agabus godmanni Crotch, 1867
Agabus granulatus (Falkenström, 1936)
Agabus gringo Larson, 2000
Agabus griseipennis LeConte, 1859
Agabus guttatus (Paykull, 1798)
Agabus heydeni Wehncke, 1872
Agabus hoppingi Leech, 1942
Agabus hummeli (Falkenström, 1936)
Agabus immaturus Larson, 1991
Agabus inexspectatus Nilsson, 1990
Agabus infuscatus Aubé, 1838
Agabus inscriptus (Crotch, 1873)
Agabus jacobsoni Zaitzev, 1905
Agabus japonicus Sharp, 1873
Agabus kaszabi Guéorguiev, 1972
Agabus kholini Nilsson, 1994
Agabus klamathensis Larson & Leech, 1989
Agabus kokoosson Feng, 1936
Agabus kootenai Larson, 1991
Agabus labiatus (Brahm, 1790)
Agabus laferi Nilsson, 1994
Agabus lagabrunensis Schizzerotto & Fery, 1989
Agabus lapponicus (Thomson, 1867)
Agabus leptapsis (LeConte, 1878)
Agabus lineatus Gebler, 1848
Agabus lithax Riha, 1961
Agabus lobonyx Guignot, 1952
Agabus loeffleri Wewalka & Nilsson, 1990
Agabus longissimus Régimbart, 1899
Agabus luteaster Zaitzev, 1906
Agabus lutosus LeConte, 1853
Agabus mackenziensis Larson, 1991
Agabus maderensis Wollaston, 1854
Agabus mandsuricus (Guignot, 1956)
Agabus margaretae Larson, 1975
Agabus matsumotoi Satô & Nilsson, 1990
Agabus melanarius Aubé, 1837
Agabus moestus (Curtis, 1835)
Agabus morosus LeConte, 1852
Agabus mucronatus (Falkenström, 1936)
Agabus nebulosus (Forster, 1771)
Agabus nevadensis Hå.Lindberg, 1939
Agabus niedzwiedzkii Lomnicki, 1894
Agabus obliteratus LeConte, 1859
Agabus oblongulus Fall, 1922
Agabus obsoletus LeConte, 1858
Agabus ommani Zaitzev, 1908
Agabus pallens Poppius, 1905
Agabus pallidus Omer-Cooper, 1931
Agabus paludosus (Fabricius, 1801)
Agabus parvulus Lomnicki, 1894
Agabus perditus Scudder, 1900
Agabus perssoni Nilsson, 1992
Agabus phaeopterus (Kirby, 1837)
Agabus philippensis (Zimmermann, 1924)
Agabus picotae Foster & Bilton, 1997
Agabus pisobius Leech, 1949
Agabus poppiusi Nilsson, 2003
Agabus pseudoclypealis Scholz, 1933
Agabus punctatus F.E.Melsheimer, 1844
Agabus punctulatus Aubé, 1838
Agabus raffrayi Sharp, 1882
Agabus ragazzii Régimbart, 1887
Agabus ramblae Millán & Ribera, 2001
Agabus rathbuni Scudder, 1900
Agabus regimbarti Zaitzev, 1906
Agabus reitteri Lomnicki, 1894
Agabus rottensis Nilsson, 2001
Agabus rufipennis (Gschwendtner, 1933)
Agabus rufulus Fairmaire, 1859
Agabus rumppi Leech, 1964
Agabus ruwenzoricus Guignot, 1936
Agabus safei Abdul-Karim & Ali, 1986
Agabus sasquatch Larson, 1991
Agabus semipunctatus (Kirby, 1837)
Agabus serricornis (Paykull, 1799)
Agabus setulosus (J.Sahlberg, 1895)
Agabus sikhotealinensis (Lafer, 1988)
Agabus sjostedti Régimbart, 1908
Agabus slovzovi (J.Sahlberg, 1880)
Agabus smithi Brown, 1930
Agabus solskii Jakovlev, 1897
Agabus strigulosus (Crotch, 1873)
Agabus striolatus (Gyllenhal, 1808)
Agabus sturmii (Gyllenhal, 1808)
Agabus subfuscatus Sharp, 1882
Agabus suoduogangi Stastný & Nilsson, 2003
Agabus svenhedini (Falkenström, 1932)
Agabus taeniolatus (Harris, 1828)
Agabus taiwanensis Nilsson & Wewalka, 1994
Agabus thomsoni (J.Sahlberg, 1871)
Agabus tibetanus Zaitzev, 1908
Agabus tristis Aubé, 1838
Agabus turcmenus Guignot, 1957
Agabus udege Nilsson, 1994
Agabus uliginosus (Linnaeus , 1761)
Agabus undulatus (Schrank, 1776)
Agabus ungeri (Heer, 1847)
Agabus unguicularis (Thomson, 1867)
Agabus uralensis Nilsson & Petrov, 2006
Agabus valdiviensis Gemminger & Harold, 1868
Agabus velox Leech, 1939
Agabus vereschaginae Angus, 1984
Agabus winkleri (Gschwendtner, 1923)
Agabus wollastoni Sharp, 1882
Agabus xyztrus Larson, 2000
Agabus yakutiae Nilsson & Larson, 1990
Agabus zetterstedti Thomson, 1856
Agabus zimmermanni Scholz, 1920
^ Zool. Misc. , 3 : 69, 72. Type species : Agabus paykullii Leach, 1817 (= Dytiscus serricornis Paykull , 1799)
^ a b Ross H. Arnett , Michael C. Thomas (2002). American Beetles: Staphyliniformi . Boca Raton, FL: CRC Press. ISBN 0849319250 . P. 175-176.
^ a b Anders N. Nilsson, Mogens Holmen (1995). The Aquatic Adephaga (Coleoptera) of Fennoscandia and Denmark: II - Dytiscidea . Leiden - a.o.: Brill. ISBN 9004104569 . P. 87.
^ Foster, G. 1996. Agabus clypealis . In: IUCN 2008. 2008 IUCN Red List of Threatened Species. Accessed on 28 November 2008.
^ Foster, G. 1996. Agabus discicollis . In: IUCN 2008. 2008 IUCN Red List of Threatened Species. Accessed on 28 November 2008.
^ Foster, G. 1996. Agabus hozgargantae . In: IUCN 2008. 2008 IUCN Red List of Threatened Species. Accessed on 28 November 2008.
^ Ignacio Ribera, Anders N. Nilsson, Alfried P. Vogler (2004). Phylogeny and historical biogeography of Agabinae diving beetles (Coleoptera) inferred from mitochondrial DNA sequences. Molecular Phylogenetics and Evolution 30 (3): 545-562.
^ Nilsson, A.N. (June 2000). "A new view on the generic classification of the Agabus-group of genera of the Agabini, aimed at solving the problem with a paraphyletic Agabus (Coleoptera: Dytiscidae)". Koleopterologische Rundschau . 70 : 17–36.
^ Dytiscidae Species List at Joel Hallan's Biology Catalog. Texas A&M University. Retrieved on 7 May 2012.