Darwinopterus (meaning "Darwin's wing") is a genus of pterosaur, discovered in China and named after biologist Charles Darwin.[1] Between 30 and 40 fossil specimens have been identified,[2] all collected from the Tiaojishan Formation, which dates to the middle Jurassic period, 160.89–160.25 Ma ago.[3][4] The type species, D. modularis, was described in February 2010.[1]D. modularis was the first known pterosaur to display features of both long-tailed (rhamphorhynchoid) and short-tailed (pterodactyloid) pterosaurs, and was described as a transitional fossil between the two groups.[5] Two additional species, D. linglongtaensis and D. robustodens, were described from the same fossil beds in December 2010 and June 2011, respectively.[6][7]
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^Liu Y.-Q. Kuang H.-W., Jiang X.-J., Peng N., Xu H. & Sun H.-Y. (2012). "Timing of the earliest known feathered dinosaurs and transitional pterosaurs older than the Jehol Biota." Palaeogeography, Palaeoclimatology, Palaeoecology (advance online publication).
^Lü J.; Xu L.; Chang H.; Zhang X. (2011). "A new darwinopterid pterosaur from the Middle Jurassic of western Liaoning, northeastern China and its ecological implications". Acta Geologica Sinica - English Edition. 85 (3): 507–514. doi:10.1111/j.1755-6724.2011.00444.x. S2CID128545851.