Immunoreceptor tyrosine-based inhibitory motif

An immunoreceptor tyrosine-based inhibitory motif (ITIM), is a conserved sequence of amino acids that is found intracellularly in the cytoplasmic domains of many inhibitory receptors of the non-catalytic tyrosine-phosphorylated receptor family found on immune cells.[1][2] These immune cells include T cells, B cells, NK cells, dendritic cells, macrophages and mast cells.  ITIMs have similar structures of S/I/V/LxYxxI/V/L, where x is any amino acid, Y is a tyrosine residue that can be phosphorylated, S is the amino acid serine, I is the amino acid isoleucine, and V is the amino acid valine.[3] ITIMs recruit SH2 domain-containing phosphatases, which inhibit cellular activation. ITIM-containing receptors often serve to target immunoreceptor tyrosine-based activation motif (ITAM)-containing receptors, resulting in an innate inhibition mechanism within cells.[3] ITIM bearing receptors have important role in regulation of immune system allowing negative regulation at different levels of the immune response.[4]

A list of human candidate ITIM-containing proteins has been generated by proteome-wide scans.[5] It has included more than 135 proteins with ITIM motif. This list is reportedly expanded by studying rare human SNPs that create the consensus signature S/I/V/LxYxxI/V/L motif.[6]

  1. ^ Dushek O, Goyette J, van der Merwe PA (November 2012). "Non-catalytic tyrosine-phosphorylated receptors". Immunological Reviews. 250 (1): 258–76. doi:10.1111/imr.12008. PMID 23046135. S2CID 1549902.
  2. ^ Veillette A, Latour S, Davidson D (2002-04-01). "Negative regulation of immunoreceptor signaling". Annual Review of Immunology. 20 (1): 669–707. doi:10.1146/annurev.immunol.20.081501.130710. PMID 11861615.
  3. ^ a b Coxon CH, Geer MJ, Senis YA (June 2017). "ITIM receptors: more than just inhibitors of platelet activation". Blood. 129 (26): 3407–3418. doi:10.1182/blood-2016-12-720185. PMC 5562394. PMID 28465343.
  4. ^ Verbrugge A, Meyaard L (June 2005). "Signaling by ITIM-bearing receptors". Current Immunology Reviews. 1 (2): 201–12. doi:10.2174/1573395054065160.
  5. ^ Staub E, Rosenthal A, Hinzmann B (April 2004). "Systematic identification of immunoreceptor tyrosine-based inhibitory motifs in the human proteome". Cellular Signalling. 16 (4): 435–56. doi:10.1016/j.cellsig.2003.08.013. PMID 14709333.
  6. ^ Ulaganathan VK (May 2020). "TraPS-VarI: Identifying genetic variants altering phosphotyrosine based signalling motifs". Scientific Reports. 10 (1): 8453. Bibcode:2020NatSR..10.8453U. doi:10.1038/s41598-020-65146-2. PMC 7242328. PMID 32439998.