Based on the examination of O. pulcher fossils from the Oligocene, the Tremoctopus–Argonauta divergence has been estimated to have occurred at least 29 million years ago.[5]
^ abcKobayashi, T. (1954). A new Palaeogene paracenoceratoid from southern Kyushu in Japan. Japanese Journal of Geology and Geography24: 181–184.
^Cite error: The named reference awaensis was invoked but never defined (see the help page).
^Ozawa, T. & S. Tomida (1996). Occurrence of Aturia coxi (Cephalopoda; Nautilida) from the uppermost Miocene of Japan and its implication for late Miocene marine climate in the northwestern Pacific. Journal of Paleontology70(5): 795–798. JSTOR1306481
^ abSaul, L.R. & C.J. Stadum (2005). Fossil argonauts (Mollusca: Cephalopoda: Octopodida) from Late Miocene siltstones of the Los Angeles Basin, California. Journal of Paleontology79(3): 520–531. doi:10.1666/0022-3360(2005)079<0520:FAMCOF>2.0.CO;2
^ abStrugnell, J., J. Jackson, A.J. Drummond & A. Cooper (2006). Divergence time estimates for major cephalopod groups: evidence from multiple genes. Cladistics22(1): 89–96. doi:10.1111/j.1096-0031.2006.00086.x
^Martill, D.M. & M.J. Barker (2006). A paper nautilus (Octopoda, Argonauta) from the Miocene Pakhna Formation of Cyprus. Palaeontology49(5): 1035–1041. doi:10.1111/j.1475-4983.2006.00578.x
^Strugnell, J.M., A.D. Rogers, P.A. Prodöhl, M.A. Collins & A.L. Allcock (2008). The thermohaline expressway: the Southern Ocean as a centre of origin for deep-sea octopuses. Cladistics24(6): 853–860. doi:10.1111/j.1096-0031.2008.00234.x
^Strugnell, J. & A.L. Allcock (2010). Co-estimation of phylogeny and divergence times of Argonautoidea using relaxed phylogenetics. Molecular Phylogenetics and Evolution54(3): 701–708. doi:10.1016/j.ympev.2009.11.017
^Ward, P. (1980). Comparative shell shape distributions in Jurassic–Cretaceous ammonites and Jurassic–Tertiary nautilids. Paleobiology6(1): 32–43. JSTOR2400233
^Teichert, C. & T. Matsumoto (2010). The Ancestry of the Genus Nautilus. In: W.B. Saunders & N.H. Landman (eds.) Nautilus: The Biology and Paleobiology of a Living Fossil. Springer. pp. 25–32. doi:10.1007/978-90-481-3299-7_2