| Phytosaurs | |
|---|---|
| |
| Skeleton of Redondasaurus at the Carnegie Museum of Natural History | |
Scientific classification 
| |
| Domain: | Eukaryota |
| Kingdom: | Animalia |
| Phylum: | Chordata |
| Class: | Reptilia |
| Clade: | Archosauromorpha |
| Clade: | Archosauriformes |
| Clade: | Eucrocopoda |
| Clade: | Crurotarsi |
| Order: | †Phytosauria von Meyer, 1861 |
| Subgroups | |
| Synonyms | |
|
Parasuchia Huxley, 1875 | |
Phytosaurs (Φυτόσαυροι in Greek, meaning 'plant lizard') are an extinct group of large, mostly semiaquatic Late Triassic archosauriform reptiles. Phytosaurs belong to the order Phytosauria and are sometimes referred to as parasuchians. Phytosauria, Parasuchia, Parasuchidae, and Phytosauridae have often been considered equivalent groupings containing the same species. Some recent studies have offered a more nuanced approach, defining Parasuchidae and Phytosauridae as nested clades within Phytosauria as a whole. The clade Phytosauria was defined by Paul Sereno in 2005 as Rutiodon carolinensis and all taxa more closely related to it than to Aetosaurus ferratus, Rauisuchus tiradentes, Prestosuchus chiniquensis, Ornithosuchus woodwardi, or Crocodylus niloticus (the Nile crocodile).[2] Phytosaurs were long-snouted and heavily armoured, bearing a remarkable resemblance to modern crocodilians in size, appearance, and lifestyle, as an example of convergence or parallel evolution.
The name "phytosaur" means "plant lizard", as the first fossils of phytosaurs were mistakenly thought to belong to plant eaters.[3]
For many years, phytosaurs were considered to be the most basal group of Pseudosuchia (crocodile-line archosaurs), meaning that they were thought to be more closely related to the crocodilians than to birds (the other living group of archosaurs). Some studies of the evolutionary relationships of early archosauriforms have suggested that phytosaurs evolved before the split between crocodile- and bird-line archosaurs and are a sister taxon of Archosauria. The most recent study retains the former way of classifying phytosaurs as pseudosuchians.[4]
Phytosaurs had a nearly global distribution during the Triassic. Fossils have been recovered from Europe, North America, India, Morocco, Thailand, Brazil, Greenland[5] and Madagascar. Fossils attributed to phytosaurs have been found in Early Jurassic rocks, possibly extending their temporal range beyond the Triassic-Jurassic boundary.[6] They may have also been present in rock layers dating to the Middle Triassic of China as evidenced by Diandongosuchus, however it is not known if this is truly a member of the clade.
- ^ "Phytosauria: The phytosaurs". University of California Museum of Paleontology. Matt Wedel. Retrieved 20 June 2022.
- ^ Nesbitt, S.J. (2011). "The early evolution of archosaurs: relationships and the origin of major clades". Bulletin of the American Museum of Natural History. 352: 1–292. doi:10.1206/352.1. hdl:2246/6112. S2CID 83493714.
- ^ Colbert, Edwin H. (Edwin Harris); Knight, Charles Robert (1951). The dinosaur book: the ruling reptiles and their relatives. New York: McGraw-Hill. p. 152.
- ^ Jones, Andrew S.; Butler, Richard J. (10 December 2018). "A new phylogenetic analysis of Phytosauria (Archosauria: Pseudosuchia) with the application of continuous and geometric morphometric character coding". PeerJ. 6: e5901. doi:10.7717/peerj.5901. PMC 6292387. PMID 30581656.
- ^ Mateus, O., Clemmensen L., Klein N., Wings O., Frobøse N., Milàn J., Adolfssen J., & Estrup E. (2014). The Late Triassic of Jameson Land revisited: new vertebrate findings and the first phytosaur from Greenland. Journal of Vertebrate Paleontology. Program and Abstracts, 2014, 182.
- ^ Maisch, M.W.; Kapitzke, M. (2010). "A presumably marine phytosaur (Reptilia: Archosauria) from the pre-planorbis beds (Hettangian) of England". Neues Jahrbuch für Geologie und Paläontologie - Abhandlungen. 257 (3): 373–379. doi:10.1127/0077-7749/2010/0076.