A secondary sex characteristic is a physical characteristic of an organism that is related to or derived from its sex, but not directly part of its reproductive system.[1] In humans, these characteristics typically start to appear during puberty—and include enlarged breasts and widened hips of females, facial hair and Adam's apples on males, and pubic hair on both.[1][2] In non-human animals, they can start to appear at sexual maturity[3]—and include, for example, the manes of male lions,[4] the bright facial and rump coloration of male mandrills, and horns in many goats and antelopes.
Secondary sex characteristics are particularly evident in the sexually dimorphic phenotypic traits that distinguish the sexes of a species.[5] In evolution, secondary sex characteristics are the product of sexual selection for traits that show fitness, giving an organism an advantage over its rivals in courtship and in aggressive interactions.[6]
Many characteristics are believed to have been established by a positive feedback loop known as the Fisherian runaway produced by the secondary characteristic in one sex and the desire for that characteristic in the other sex. Male birds and fish of many species have brighter coloration or other external ornaments. Differences in size between sexes are also considered secondary sexual characteristics.