Sphenacanthus (from Greek: σφήνsphḗn, 'wedge' and Greek: ἄκανθαakantha, 'spine')[3] is an extinct genus of a chondrichtyanxenacanthiform that belongs to the Sphenacanthidae family and lived from the Late Devonian, through Carboniferous until the Late Permian period in Scotland,[4]Spain,[5]Russia[6] and Brazil.[7] It lived 359 million years ago, and probably it was one of the first member of the elasmobranchians, the lineage that leads to the modern sharks. Sphenacanthus probably hunts small fishes and, unlike their modern-day relatives, its inhabited fresh water lagoons. Sphenacanthus had seven fins, two in the upper part and five in the underside, and it had a heterodont dentition and mandibles relatively long and deeper.[4]Sphenacanthus serrulatus is still only known from incomplete neurocranial remains and associated dermal material. These suggest that it was a relatively large shark, probably well over one meter in length when fully grown. Its body form was probably similar to that of other phalacanthous sharks.[8]
^L. Agassiz. 1837. Recherches Sur Les Poissons Fossiles. Tome III (livr. 8–9). Imprimérie de Petitpierre, Neuchatel viii-72
^Soler-Gijón, R. 1997. Euselachian sharks from the Late Carboniferous of the Puertollano Basin, Spain: bioestratigraphicand palaeoenvironmental implications. Modern Geology, 21:137–169.
^A. Ivanov. 1999. Late Devonian – Early Permian chondrichthyans of the Russian Arctic. Acta Palaeontologica Polonica 49(3):267–285
^JOHN R. F. DICK, Sphenacanthus, a Palaeozoic freshwater shark, Zoological Journal of the Linnean Society, Volume 122, Issue 1–2, January 1998, Pages 9–25, doi:10.1111/j.1096-3642.1998.tb02523.x