Stygofauna

Stygofauna are any fauna that live in groundwater systems or aquifers, such as caves, fissures and vugs. Stygofauna and troglofauna are the two types of subterranean fauna (based on life-history). Both are associated with subterranean environments – stygofauna are associated with water, and troglofauna with caves and spaces above the water table. Stygofauna can live within freshwater aquifers and within the pore spaces of limestone, calcrete or laterite, whilst larger animals can be found in cave waters and wells. Stygofaunal animals, like troglofauna, are divided into three groups based on their life history - stygophiles, stygoxenes, and stygobites.

1. Stygophiles inhabit both surface and subterranean aquatic environments, but are not necessarily restricted to either.
2. Stygoxenes are like stygophiles, except they are defined as accidental or occasional presence in subterranean waters. Stygophiles and stygoxenes may live for part of their lives in caves, but don't complete their life cycle in them.
3. Stygobites are obligate, or strictly subterranean, aquatic animals and complete their entire life in this environment.^[1]

Extensive research of stygofauna has been undertaken in countries with ready access to caves and wells such as France, Slovenia, the US and, more recently, Australia. Many species of stygofauna, particularly obligate stygobites, are endemic to specific regions or even individual caves. This makes them an important focus for the conservation of groundwater systems.