Thymic involution

Thymic involution is the shrinking (involution) of the thymus with age, resulting in changes in the architecture of the thymus and a decrease in tissue mass.[1] Thymus involution is one of the major characteristics of vertebrate immunology, and occurs in almost all vertebrates, from birds, teleosts, amphibians to reptiles, though the thymi of a few species of sharks are known not to involute.[1][2] This process is genetically regulated, with the nucleic material responsible being an example of a conserved sequence — one maintained through natural selection (though the pressures shaping this are unclear as will be discussed) since it arose in a common ancestor of all species now exhibiting it, via a phenomenon known to bioinformaticists as an orthologic sequence homology.

T-cells are named for the thymus where T-lymphocytes migrate from the bone marrow to mature. Its regression has been linked to the reduction in immunosurveillance[3] and the rise of infectious disease and cancer incidence in the elderly (in some cases risk is inversely proportional to thymus size).[4] Though thymic involution has been linked to immunosenescence, it is not induced by senescence as the organ starts involuting from a young age:[5] in humans, as early as the first year after birth.[6]

  1. ^ a b Shanley D.P.; Danielle A.W.; Manley N.R.; Palmer D.B.; et al. (2009). "An evolutionary perspective on the mechanisms of immunosenescence" (PDF). Trends in Immunology. 30 (7): 374–381. doi:10.1016/j.it.2009.05.001. PMID 19541538.
  2. ^ Zakharova L.A. (2009). "Evolution of adaptive immunity". Izvestiia Rossiĭskoĭ Akademii Nauk. Seriia Biologicheskaia. 2 (2): 143–154. PMID 19391473.
  3. ^ Linton P.J.; Dorshkind K. (2004). "Age-related changes in lymphocyte development and function". Nature Immunology. 5 (2): 133–139. doi:10.1038/ni1033. PMID 14749784. S2CID 12485241.
  4. ^ Palmer S.; Albergante L.; Blackburn C.C.; Newman T.J. (2018). "Thymic involution and rising disease incidence with age". Proceedings of the National Academy of Sciences of the United States of America. 115 (8): 1883–1888. Bibcode:2018PNAS..115.1883P. doi:10.1073/pnas.1714478115. PMC 5828591. PMID 29432166.
  5. ^ Taub D.D.; Long D.L. (2005). "Insights into thymic aging and regeneration". Immunological Reviews. 205: 72–93. doi:10.1111/j.0105-2896.2005.00275.x. PMID 15882346. S2CID 24461464.
  6. ^ Steinmann G.G.; Klaus B.; Muller-Hermelin H.K.; et al. (1985). "The involution of the aging human thymic epithelium is independent of puberty. A morphometric study". Scandinavian Journal of Immunology. 22 (5): 563–75. doi:10.1111/j.1365-3083.1985.tb01916.x. PMID 4081647. S2CID 40226062.